Botany & Zoology
Welcome to Stellenbosch University

Prof. Savel Daniels

​​Evolutionary Genomics Group

Research focus

One of the central focal points of my research is to understand the interplay between species diversity and historical time in southern Africa. In this respect I have focused on evolutionary relationships of a wide number of terrestrial and inland aquatic groups, primarily invertebrates. In this respect I am currently investigating the systematics of South African velvet worms and a revision of the Afrotropical freshwater crab fauna based on DNA sequence data and morphology. I have also in recent years worked extensively on limbless skinks and have recently initiated a large-scale project on tortoises. The main objectives of all these studies are to better understand and document the spectacular biological diversity of South Africa to aid the conservation of these groups.

Current collaboration

South Africa

Dr Michelle Hamer, SANBI, Pretoria (velvet worm systematics)

Prof Neil Heideman, Dean, University of the Free State, Bloemfontein (Systematics of limbless lizards)

Dr Mike Picker, Zoology Department, University of Cape Town, Rondebosch, Cape Town (comparative phylogeography of selected mountain invertebrate species)

Dr Gavin Gouws, SAIAB, Grahamstown, South Africa (aquatic biodiversity)

Prof Retha Hofmeyr, Department of Biodiversity and Conservation, University of the Western Cape, Bellville, South Africa (tortoise phylogeography and systematics)


Prof Hilke Ruhberg, Biozentrum Grindel und Zoologisches Museum, Hamburg, Germany (velvet worm systematics)


Prof Barbara Stewart, Centre of Excellence in Natural Resource Management, University of Western Australia, Australia (comparative phylogeography of selected aquatic invertebrate species)


Dr Trisha Spears, Department of Biological Sciences, Florida State University, Tallahassee, Florida, U.S.A. (Phylogenetics of the Decapod Crustacea)

Prof Keith Crandall, Brighman Young University, Department of Integrative Biology, Provo, Utah, USA
(Systematics of freshwater decapods)

Prof Neil Cumberlidge, Department of Biology, Northern Michigan University, Presque Isle Ave, Marquette, USA (Systematics of freshwater crabs)

Prof Gonzalo Giribet, Museum of Comparative Zoology, Harvard University, Cambridge, USA (velvet worm phylogenetics)


99) Busschau, T., Conradie, W., Daniels, S.R. (2021). One species hides many: molecular and morphological evidence for cryptic speciation in a thread snake (Leptotypholopidae: Leptotyphlops sylvicolus). Journal of Zoological Systematics and Evolutionary Research, 59: 195-221.

98) Raphalo, E.M., Cole, M.L., Daniels, S.R. (2021). Climatic oscillations during the Mio/Pliocene epoch induced cladogenesis in the terrestrial snail genus Gittenedouardia (Mollusca: Gastropoda: Cerastidae) from South Africa. Molecular Phylogenetics and Evolution, 155: 107000.

97) Daniels, S.R., James, N.C., Gouws, G. (2020). Phylogeographic structure and continued surveys of a vulnerable South African freshwater crab (Potamonautidae: Potamonautes lividus) implications for the IUCN Red Listing of the Afrotropical fauna. Aquatic Conservation: Marine and Freshwater ecosystems, 30: 2221-2239.

96) Barnes, A., Reiss, T., Daniels, S.R. (2020). Systematics of the Peripatopsis clavigera species complex (Onychophora: Peripatopsidae) reveals cryptic cladogenic patterning, with the description of five new species. Invertebrate Systematics, 34: 569-590.

95) Daniels, S.R., Bittencourt-Silva, G.B., Muianga, V., Bayliss. J. (2020). Phylogenetics of the freshwater crab (Potamonautes MacLeay, 1838) fauna from 'sky islands' in Mozambique with the description of a new species. European Journal of Taxonomy, 716: 1-23.

94) Cumberlidge, N., Soma, J.B., Leever, E.M., Daniels, S.R. (2020). New lineages in the Malagasy freshwater crab fauna: Agora n. gen. for Thelphusa goudi H. Milne Edwards, 1853, and a phytotelmic new species of Malagasya. Journal of Crustacean Biology, 40: 584-599.

93) Kushata, J.N.T., Conradie, W., Cherry, M.I., Daniels, S.R. (2020). Comparison of the mitochondrial phylogeographic structure of a generalist and two specialist forest species reveals contrasting patterns in the Eastern and Western Cape provinces of South Africa. Biological Journal of the Linnean Society, 130: 783-799.

92) Hofmeyr, M.D., Ihlow, F., Fouche, P., Daniels, S.R. (2020).  Niche divergence corresponds to genetic differentiation within the parrot-beak tortoise, Homopus areolatus, endemic to South Africa. Zoological Journal of the Linnean Society, 190: 1256-1273.

91) Kochey, J. K., Daniels, S.R., Plagge, C., Mebrabi, S., Hartmann, L, Schrenk, F., Plath, M., Klaus, S. (2019). Genetic differentiation of the Malawi blue crab reflects Pleistocene desiccation of Lake Malwi (Brachyura, Potamonautidae: Potamonautes lirrangensis (Rathbun, 1904). Hydrobiologia, 843: 1-11.

90) Busschau, T., Conradie, W., Daniels, S.R. (2019). Evidence for cryptic diversification in a rupicolous forest-dwelling gecko (Gekkonidae: Afroedura pondolia) from a biodiversity hotspot. Molecular Phylogenetics and Evolution, 139: 106549.

89) Daniels, S.R., Busschau, T., Cumberlidge, N. (2019). Two new species of freshwater crabs of the genus Potamonautes MacLeay, 1838 (Decapoda: Brachyura: Potamonautidae). Journal of Crustacean Biology, 39: 426-435.

88) Gibson, K.M., Nguyen, B.N., Neumann, L.M., Miller, M., Buss, P., Daniels, S., Ahn, M.J., Crandall, K.A., Pukazhenthi, B. (2019). Gut microbiome differences between wild and captive black rhinoceros – implications for rhino health. Scientific Reports, 9: 1-11

87) Cumberlidge, N., Ndongo, PAM, Clark, P.F., Daniels, S.R. (2019). A new genus for the freshwater crab Potamonemus asylus Cumberlidge, 19993 (Brachyura: Potamoidea: Potamonautidae) from Cameroon, Central Africa, with a key to the genera in the Potamonautidae. Journal of Natural History, 53: 659-676.

86) Barnes A., Daniels, S. R. (2019). On the importance of fine-scale sampling in detecting alpha taxonomic diversity among saproxylic invertebrates: a velvet worm template (Onychophora: Opisthopatus amaxhosa). Zoologica Scripta, 48: 243-262.

85) Wood, L. E., de Grave, S., Cumberlidge, N., Kennerley, A., Daniels, S. R. (2019). A phylogenetic perspective on biodiversity patterns of Afrotropical freshwater crabs (Crustacea, Decapoda, Potamonautidae). Aquatic conservation: of Freshwater and Marine ecosystems, 29: 2219-2230.

84) Kullenkampff, K., van Zyl, F., Klaus, S., Daniels, S. R. (2019). Molecular evidence for cryptic species in the common slug eating snake, Duberria lutrix lutrix from South Africa. ZooKeys, 838: 133-154.

83) Wood, L. E., de Grave, S., Daniels, S. R. (2019). A comparative evolutionary study reveals radically different scales of genetic structure in two atyid shrimp species. Zoological Journal of the Linnean Society, 186: 200-212.

82) Cumberlidge, N., Daniels, S.R. (2018). A new species of Potamonautes from São Tomé Island, Central Africa, with the redescription of P. margaritarius from São Tomé and P. principe from Príncipe (Decapoda: Potamonautidae). Contributions to Zoology, 87: 287-303.

81) Giribet, G., Buckman-Young, B., Costa, CS., Baker, C. M., Benavides L. R., Bransterrer, M. G., Daniels, S.R., Pinto-da-Rocha, R.  (2018) The 'Peripatos' in Eurogondwana? – Lack of evidence that south-east Asian onychophorans walked through Europe. Invertebrate Systematics, 32: 842-865.

80) Nielson, S. V., Daniels, S.R., Conradie, W., Heinicke, M, P., Noonan. B. P. (2018). Multilocus phylogenetics in a widespread African anuran lineage (Breviciptidae: Breviceps) reveals patterns of diversity reflecting geoclimatic changes. Journal of Biogeography, 45: 2067-2079.

79) Gouws, G., Daniels, S. R., MacDonald A. H., Chakona A and Barker, NP. (2018). Development and characterization of a microsatellite library for the freshwater crab Potamonautes danielsi and its transferability across three congeneric species. Journal of Crustacean Biology, 38: 765-771.

78) Daniels, S. R., Klaus, S. (2018). Divergent evolutionary origins and biogeographic histories of two freshwater crabs (Brachyura: Potamonautes) on the West African conveyer belt islands of São Tomé and Príncipe. Molecular Phylogenetics and Evolution, 129: 119-128.

77) Wood, L. E., Daniels, S. R., De Drave, S. (2018). Caridina susorboflora is a junior synonym of the widespread C. africana. Crustaceana, 91: 243-249.

76) Ng, P., Daniels, S. R. (2018). A new species of the trogloplacine crab of the genus Australocarcinus Davies, 1988 from a freshwater stream on Mahe, Seychelles. ZooKeys, 738: 27-35.

75) Daniels, S. R., Dreyer, M., Sharma, P. (2017). Contrasting the population genetic of two velvet worm taxa (Onychophora: Peripatopsidae: Peripatopsus) in forest fragments along the south-eastern Cape, South Africa.  Invertebrate Systematics 31: 781-796.

74) Wood, L. E., de Grave S., van Heerden C., Daniels, S.R. 2017. Complete mitochondrial genome of the freshwater prawn Paleomon capensis (Crustacea: Palaemonidae). Mitochondrial DNA (B), 2: 742-743.

73) Daniels, S. R. 2017. Sympatric colour morphs or distinct taxa: Examining species boundaries among two south African freshwater crabs with the description of a new species. Journal of Crustacean Biology, 37: 723-731.

72) Hofmeyr, M.D., Vamberg, M., Branch, W., Schleicher A., Daniels, S.R.  2017. Tortoise (Reptilia, Testudinidae) radiations in Southern Africa from the Eocene to the present. Zoological Scripta, 46: 389-400.

71) Busschau, T., Conradie, W., Jordaan, A., Daniels, S. R. 2017. Unmasking evolutionary diversity among two closely related South African legless skink species (Acontinae: Acontias) using molecular data. Zoology, 121: 72-82.

70) Wood, L. E., de Grave S, Daniels, S. R. 2017.  Phylogeographic patterning among two co-distributed shrimp species (Crustacea: Decapoda: Palaemonidae) reveal high levels of connectivity across biogeographic regions along the South African coast. PLoS-one 12: e0173356

69) Wood, L. E and Daniels, S. R. 2016. Genetic and morphological evidence for a new mountain-living freshwater crab species (Decapoda: Potamonautidae: Potamonautes) from the Western Cape province of South Africa. Invertebrate Systematics, 30: 219-230.

68) Phiri, E. E and Daniels, S. R. 2016. Multilocus coalescent species delimitation

reveals widespread cryptic differentiation among Drakensberg mountain living

freshwater crabs (Decapoda: Potamonautes). Invertebrate Systematics, 30: 60-74.

67) Giribet, G., Boyer, Baker, S. L., Fernández, R., Sharma, P. P., de Bivort, B. L., Daniels, S.R., Harvey M, S and Griswold, C. E. 2016. A molecular phylogeny of the temperate Gondwanan family Pettalidae (Arachnida, Opiliones, Cyphophthalmi) and the limits of taxonomic sampling. Zoological Journal of the Linnean Society, 178: 523-545.

66) Daniels, S. R., Damberi, C.,Klaus, S., Sharma, P. P. (2016) Unmasking alpha diversity, cladogenesis and biogeographical patterning in an ancient panarthropod lineage (Onychophora: Peripatopsidae: Opisthopatus cinctipes) with the description of five novel species. Cladistics, 32: 506-537.

65) Daniels, S. R., McLeod, C,  Carveth, C., Mexim, K. K, and Cumberlidge, N. (2016). Examining the evolutionary relationships amongst three species of West African freshwater crabs Liberonautes Bott, 1955 (Brachyura: Potamonautidae) using mtDNA sequence data. Journal of Crustacean Biology, 36: 731-739.

64) Daniels, S.R., Phiri E.E., Klaus S., Albrecht C., Cumberlidge N. (2015). Multi-locus phylogeny of the Afrotropical freshwater crab fauna reveals historical drainage connectivity and transoceanic dispersal since the Eocene. Systematic Biology, 64: 549-567.

63) Mirimin, L., Kitchin N., Impson, D. N., Clark, P. F., Richard, J., Daniels, S.R., Roodt-Wilding, R. (2015).  Genetic and morphological characterization of freshwater shrimps (Caridina africana Kingsley, 1882) reveals the presence of alien shrimp in the Cape Floristic Region, South Africa. Journal of Heredity, 106: 711-718. 

62) Myburg, A., Daniels, S.R. (2015). Exploring the impact of habitat size on phylogeographic patterning in the Overberg velvet worm Peripatopsis overbergiensis (Onychophora: Peripatopsidae). Journal of Heredity, 106: 296-305.

61) Daniels, S.R., Phiri, E. E., Bayliss J. (2014). Renewed sampling of inland aquatic habitats in southern Africa yields two novel freshwater crab species (Decapoda: Potamonautidae: Potamonautes). Zoological Journal of the Linnean Society, 171: 356-369.

60) Diedericks, G., and Daniels, S.R. (2014). Ain't no mountain high enough, ain't no valley low enough? Phylogeography of the rupicolous Cape girdled lizard (Cordylus cordulys). Molecular Phylogenetics and Evolution, 71: 234-248.

59) Phiri, E.E., and Daniels S.R. (2014).  Disentangling the divergence and cladogenesis in the freshwater crab species (Potamonautidae: Potamonautes perlatus sensu lato) in the Cape Fold Mountains, South Africa, with the description of two novel cryptic lineages. Zoological Journal of the Linnean Society, 170: 310-332.

58) Murien, J., Daniels, S.R., Buckley, T., Mayer, G., and Giribet. G. A living fossil tale of Pangaean biogeography. Proceedings of the Royal Society of London, Biological Sciences, 281: 1775 (20132648).

57) Cumberlidge, N., and Daniels, S.R. (2014). Recognition of two new species of freshwater crabs from the Seychelles based on molecular evidence (Potamoidea: Potamonautidae).  Invertebrate Systematics, 24: 384-356.

56) Phirie, E.E., and Daniels, S.R. (2013). Hidden in the highlands: the description and phylogenetic position of a novel endemic freshwater crab from Zimbabwe. Invertebrate Systematics, 27: 530-539.

55) Daniels, S.R., McDonald, D.E., Picker, M. (2013). Evolutionary insight into the Peripatopsis balfouri sensu lato species complex (Onychophora: Peripatopsidae) reveals novel lineages and zoogeographic patterning. Zoologica Scripta, 42: 656-674.

54) Ruhberg, H and Daniels, S.R. (2013). Morphological assessment supports the recognition of four novel species in the widely distributed velvet worm Peripatopsis moseleyi (Onychophora: Peripatopsidae). Invertebrate Systematics, 27: 131-145.

53) Engelbrecht, H., Heideman, N.J.L., van Niekerk, A and Daniels, S.R. (2013).  Tracking the impact of Pliocene/Pleistocene sea level and climatic oscillations on the cladogenesis of the Cape legless fossorial skink, Acontias meleagris meleagris. Journal of Biogeography, 40: 492-506.

52) Weldon, C., Daniels, S.R., Clusella-Trullas, S and Chown, S.L. (2013). Metabolic and water loss rates of two cryptic species in the South African velvet worm Opisthopatus cinctipes (Onycophora). Journal of Comparative Physiology –B, 183: 323-332.

51) McDonald, D.E., and Daniels, S.R. (2012). Two new velvet worm species from the Western Cape, South Africa. Zootaxa, 3380: 55-68.

50) Daniels, S.R., and Bayliss, J. (2012). Neglected refugia of biodiversity: Mountainous regions in Mozambique and Malawi yield three novel freshwater crab species (Potamonautidae: Potamonautes). Zoological Journal of the Linnean Society, 164: 498-509.

49) McDonald, D.E., and Daniels, S.R. (2012). Phylogeography of the Cape velvet worm (Onycophora: Peripatopsis capensis) reveals the impact of Plio/Pleistocene climatic oscillations on Afromontane forest in the Western Cape, South Africa. Journal of Evolutionary Biology, 25: 824-835.

48) Daniels, S.R. (2011). Genetic variation in the Critically endangered velvet worm Opisthopatus roseus (Onycophora: Peripatopsidae). African Zoology 46: 419-424.

47) Daniels, S.R. (2011). Reconstructing the colonisation and diversification history of the endemic freshwater crab (Seychellum alluaudi) in the granitic and volcanic Seychelles. Molecular Phylogenetics and Evolution, 61: 534-542.

46) Heideman, N.J.L., Mulcahy, D., Sites. J. W., Hendricks, M.G and Daniels, S.R. (2011). Cryptic diversity and morphological convergence in threatened species of fossorial skinks in the genus Scelotes (Squamata: Scincidae) from the Western Cape Coast of South Africa: Implications for species boundaries, digit reduction and conservation. Molecular Phylogenetics and Evolution, 61: 823-833.

45) Voua Otomo, P., Owojori, Reinceke, S.A, Daniels, S., Reinecke, A. J. (2011). Using estimates of metal bioavailability in the soil and genetic variation of allozymes to investigate heavy metal tolerance in the earthworm Eisenia fetida. Ecotoxicology and Environmental Safety, 74: 2070-2074.

44) Engelbrecht, A, Daniels, S.R., Taylor, P.J and Rambau, R.V. (2011). Cryptic speciation in the southern African vlei rat, Otomys irroratus complex: evidence derived from mitochondrial cyt b and niche modeling.  Biological Journal of the Linnean Society, 104: 192-206.

43) Engelbrecht, A, Taylor, P.J, Daniels, S.R and Rambau, R.V. (2011). Chromosomal polymorphisms in the African vlei rats, Otomys irroratus detected using banding techniques and chromosome painting: inversions, centromeric shifts, and diploid number variation. Cytogenetics and Genome Research, 133: 8-15.

42) Engelbrecht, H. M., Mouton, P. le Fras N. and Daniels, S.R. (2011). Are melanistic populations of the Karoo girdled lizard, Karusasaurus polyzonus, relics or ecotypes? A molecular investigation. African Zoology, 46:146-155.

41) Daniels, S.R., Hofmeyr, M.D., Henen, B.T., Baard, E.H.W. (2010). Systematics and phylogeography of a threatened tortoise, the speckled padloper. Animal Conservation, 13: 237-246.

40) Allwood, J., Gleeson, D., Mayer, G., Daniels, S., Begg, J.R and Buckley, T.R. (2010). Support for vicarant origin of the New Zealand Onychophora. Journal of Biogeography, 37: 669-681.

39) Fritz, U., Daniels, S.R., Hofmeyr, M.D., Gonzalez, J., Barrio-Amoros, C.L., Siroky, P., Handsdorfer, A.K., Stukas, H. (2010). Mitochondrial phylogeography and subspecies of the wide-ranging sub-Saharan leopard tortoise Stigmochelys pardalis (Testudines: Testudinidae) – a case study for the pitfalls of pseudogenes and GenBank sequences. Journal of Zoological Systematics and Evolutionary Research, 48: 348 -359. 

38) Gouws, G., Stewart, B.A., Daniels, S.R. (2010) Phylogeographic in the gilgie (Decapoda: Parastacidae: Cherax quinquecarinatus) – a south-western Australian freshwater crayfish. Biological Journal of the Linnean Society, 101: 385-402.

37) Vargas, M., Vences, M., Branch, W. R., Daniels, S.R., Glaw, F., Hofmeyr, M.D., Kuchling, G., Maran, J., Papenfuss, T.J., Siroky, P., Vieites, D.R., Fritz, U. (2010). The widely distributed African helmeted terrapin represents a species complex: evidence from mitochondrial and nuclear DNA (Testudines: Pelomedusidae: Pelomedusa subrufa. Molecular Phylogenetics and Evolution, 56: 428-440.

36) Braband, A., Podsiadlowski, L., Cameron, S.L., Daniels, S.R., Mayer, G. (2010a). Extensive duplication events account for multiple control regions and pseudogenes in the mitochondrial genome of the velvet worm Metaperipatus inae (Onychophora, Peripatopsidae). Molecular Phylogenetics and Evolution, 57: 293-300.

35) Braband, A., Cameron, S.L., Podsiadlowski, L., Daniels, S.R., Mayer, G. (2010b). The mitochondrial genome of the onychophoran Opisthopatus cinctipes (Peripatopsidae) reflects the ancestral mitochondrial gene arrangement of Panarthropoda and Ecdysozoa. Molecular Phylogenetics and Evolution, 57:285-292.

34) Daniels, S.R., Ruhberg, H. (2010). Molecular and morphological variation in a South African velvet worm, Peripatopsis moseleyi (Onychophora, Peripatopsidae): evidence for cryptic speciation. Journal of Zoology, 282: 171-179.

33) Daniels, S.R., Picker, M., Cowlin, R.M., Hamer, M. (2009). Unravelling evolutionary lineages among the South African velvet worms (Onychophora: Peripatopsis) provides evidence for widespread cryptic speciation. Biological Journal of the Linnean Society, 97: 200-216.

32) Daniels, S.R., Heideman, N.J., Hendricks, M.G. (2009). Examining the evolutionary relationships in the Cape fossorial skink species complex (Acontinae: Acontias meleagris meleagris) reveals the presence of five cryptic lineages. Zoologica Scripta, 38: 449-463.

31) Cumberlidge, N., Ng, P.K.L., Yeo, D.C.J., Magalhaes, C., Campos, R.M., Alvarez, F., Naruse, T., Daniels, S.R., Esser, L.J., Felix, C., Attipoe, K., Clotilde-Ba, F., Darwall, W., McIvor, A., Ram, M. (2009) Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation, 142: 1665-1673.

30) Cumberlidge, N., Daniels, S.R. (2008). A conservation assessment of the freshwater crabs of southern Africa (Brachyura: Potamonautidae). African Journal of Ecology, 46: 74-79.

29) Yeo, D.C.J., Ng, P.K.L., Cumberlidge, N., Magalhaes, C., Daniels, S.R., Campos, M.R. (2008). A global assessment of freshwater crab diversity (Crustacea: Decapoda: Brachyura). Hydrobiologia, 595: 275-286.

28) Cumberlidge, N., Von Sternberg, R., Daniels, S.R. (2008). A revision of the higher taxonomy of the Afrotropical freshwater crabs (Decapoda: Brachyura) with a discussion of their biogeography. Biological Journal of the Linnean Society, 93: 399-413.

27) Mortimer, E., McGoech, M.A., Daniels, S.R., Jansen van Vuuren, B. (2008). Growth form and population genetic structure of Azorella selago on sub-Antarctic Marion Island. Antarctic Science, 20: 381-390.

26) Heideman, N.J.L., Daniels, S.R., Hendricks, M.G.J. (2008) Maximizing empirical data in delimiting species boundaries: a morphological based discriminant function analysis of the limbless skink subfamily Acontinae. African Zoology, 43: 192-201.

25) Myburgh, M., Chown, S.L., Daniels, S.R., Jansen van Vuurnen, B. (2007). Population structure, propagule pressure and conservation biogeography in sub-Antarctic: lesions from indigenous and invasive springtails. Diversity and Distribution, 13: 143-154.

24) Daniels, S.R., Hofmeyr, M.D., Henen, B.T., Crandall, K.A. (2007).  Living with the genetic signature of Miocene induced changes: evidence from the phylogeographic structure of the endemic angulate tortoise Chersina angulata. Molecular Phylogenetics and Evolution, 45: 915-926.

23) Daniels, S.R., Cumberlidge, N., Perez-Losada, M., Crandall, K.A. (2006). Evolution of Afrotropical freshwater crab lineages obscured by morphological convergence. Molecular Phylogenetics and Evolution, 40: 227-235.

22) Daniels, S.R., Gouws, G., Crandall, K.A. (2006). Phylogeographic patterning in a freshwater crab species (Decapoda: Potamonautidae: Potamonautes) reveals the signature of historic climatic oscillations. Journal of Biogeography, 33: 1538-1549.

21) Marijnissen, S.A., Daniels, S.R., Schram. F., Michelle, E. (2006). Rapid recent divergence of freshwater crabs (Platythelphusidae: Platythelphusa). Molecular Phylogenetics and Evolution, 40: 628-634.

20) Gouws, G., Stewart, B.A., Daniels, S.R. (2006). Phylogeography of the koonac (Decapoda: Parastacidae: Cherax preissii) in Western Australia: multiple lineages but only one species. Marine and Freshwater Research, 57: 837-848.

19) Daniels, S.R., Heideman, N., Hendricks, M., Crandall, K.A. (2006) Taxonomic subdivisions within the fossorial skink subfamily Acontinae (Sauria: Scincidae:) reconsidered: a multi-locus perspective. Zoologica Scripta, 35: 353-362.

18) Daniels, S.R., Heideman, N., Hendricks, M., Mokone, M.E., Crandall, K.A. (2005). Unraveling evolutionary lineages the fossorial skink group Acontias: are subspecies equivalent systematic units. Molecular Phylogenetics and Evolution, 34: 645-654.

17) Stewart, B.A., Gouws, G, Matthee, C.A., Daniels, S.R. (2004). Delimitations of cryptic sponge crab species on the genus Pseudodromia from South Africa. Zoological Scripta, 33: 45-55.

16) Gouws, G., Stewart, B.A., Daniels, S.R. (2004). Cryptic species within the freshwater isopod Mesamphisopus capensis in the Western Cape, South Africa: allozyme, morphometrics and 12 S rRNA sequence data. Biological Journal of the Linnean Society, 81: 235-253.

15) Daniels, S.R., Hamer, M., Rogers, D.C. (2004). Molecular evidence suggest an ancient radiation for the fairy shrimp genus Streptocephalus (Branchiopoda: Anostraca). Biological Journal of the Linnean Society, 82: 313-327.

14) Daniels, S. R., Mouton, P.le F.N., du Toit D. (2004). Molecular data suggest that melanistic ectotherms at the southwestern tip of Africa are the products of Miocene climatic events: evidence from cordylid lizards. Journal of Zoology, London, 263: 373-383.

13) Daniels, S.R., Gouws, G., Stewart, B. A., Coke, M. (2003).  Molecular and morphometric evidence for the presence of cryptic lineages among freshwater crabs from the Drakensberg Mountains, South Africa. Biological Journal of the Linnean Society, 78: 129-147.

12) Daniels, S.R. (2003). Examining the genetic structure among populations of the common Cape River crab, Potamonautes perlatus from river systems in South Africa reveals hydrographic boundaries. Journal of Crustacean Biology, 24: 936-950.

11) Gouws, G., Daniels, S.R., Stewart, B.A. (2002). Allozyme electrophoresis demonstrates the presence of a species boundary in freshwater crabs (Decapoda: Potamonautidae). Journal of Natural History, 36: 1199-1222.

10) Daniels, S.R., Stewart, B.A., Cook, P.A (2002). Congruent patterns of genetic variation in a burrowing freshwater crab revealed by allozyme and mt DNA sequence analysis. Hydrobiologia, 468: 171-179.

9) Daniels, S.R., Heideman, N., Hendricks, M., Wilson, B. (2002). A molecular phylogeny of the South African limbless lizard taxa of the subfamily Acontinae with special emphasis on the relationships within the genus Acontias. Molecular Phylogenetics and Evolution, 24: 314 -322.

8) Daniels, S.R., Stewart, B.A., Gouws, G., Cunningham, M., Matthee, C.A. (2002). Phylogenetic relationships of the southern African freshwater crab fauna derived from multiple data sets reveal biogeographic patterning. Molecular Phylogenetics and Evolution, 25: 511-523.

7) Daniels, S.R., Stewart, B.A., Ridgeway, T., Florance, W. (2001). Carapace dentition patterns, morphometrics and genetic differentiation amongst two toothed river crab species (Potamonautes warreni and P. unispinus) from river systems in South Africa. Journal of Zoology, London, 255: 389-404.

6) Daniels, S.R. (2001). Allometric growth, handedness and morphological variation, with a redescription of Potamonautes warreni (Calman, 1918). Crustaceana, 74: 237-253.

5) Daniels, S.R., Stewart, B.A., Burmeister, L. (2001). Geographic patterns of genetic and morphological divergence amongst populations of a freshwater crab with the description of a species from mountain streams in the Western Cape. Zoologica Scripta, 30: 181-197.

4) Daniels, S.R., Stewart, B.A., Gibbons, M.J. (1999). Genetic differentiation amongst populations of Potamonautes perlatus in the Olifants River system. Journal of Zoology, London, 249: 137-142.

3) Daniels, S.R., Stewart, B.A., Gibbons, M.J. (1999). Allozyme variation amongst populations of Potamonautes perlatus from the Berg River system. South African Journal of Zoology, 34: 64-68.

2) Daniels, S.R., Stewart, B.A., Gibbons, M.J. (1998). Genetic and morphometric variation amongst populations of Potamonautes parvispina from two river systems in the Western Cape. Journal of Natural History, 32: 1245-1258.

1) Daniels, S.R., Stewart, B.A., Gibbons, M.J. (1998). Potamonautes granularis sp. nov, a new cryptic species of river crab from the Olifants River System, South Africa. Crustaceana, 71: 885-903.​